Abstract: Tropical cyclones can form from many different precursors, including baroclinic systems. The process of an extratropical system evolving into a warm core tropical cyclone is defined by Davis and Bosart (2004) as a Tropical Transition (TT) with further classification of systems into Weak Extratropical Cylclones (WEC) and Strong Extratropical Cyclones (SEC). It is difficult to predict which systems will make the transition and which will not, but the description of a common type of TT occurring along a front will aid forecasters in identifying systems that might undergo TT. A wind speed and SST relationship thought to be necessary for this type of transition is discussed. QuikSCAT and other satellite data are used to locate TT cases forming along fronts and track their transformation into tropical systems. Frontal TT is identified as a subset of SEC TT and the evolution from a frontal wave to a tropical system is described in five stages. A frontal wave with stronger northerly wind and weaker southerly wind is the first stage in the frontal cyclogenesis. As the extratropical cyclogenesis continues in the next two stages, bent back warm front stage and instant occlusion stage, the warmer air of the bent back front becomes surrounded by cooler air . Next, in the subtropical stage the latent heat release energy from the ocean surface begins ascent and forms a shallow warm core. As the energy from surface heat fluxes translates to convection within the system, the warm core extends further into the upper levels of the atmosphere in the final, tropical stage of TT. Model data from MM5 simulations of three storms, Noel (2001), Peter (2003) and Gaston (2004) are analyzed to illustrate the five stages of frontal TT. Noel is found to have the most baroclinic origin of the three and Gaston the least.

Abstract: Advances in in situ imaging allow enumeration of abundant populations of large Rhizarians that compose a substantial proportion of total mesozooplankton biovolume. Using a quasi-Lagrangian sampling scheme, we quantified the abundance, vertical distributions, and sinking&#8208;related mortality of Aulosphaeridae, an abundant family of Phaeodaria in the California Current Ecosystem. Inter&#8208;cruise variability was high, with average concentrations at the depth of maximum abundance ranging from < 10 to > 300 cells m&#8722;3, with seasonal and interannual variability associated with temperature&#8208;preferences and regional shoaling of the 10°C isotherm. Vertical profiles showed that these organisms were consistently most abundant at 100&#65533;150&#8201;m depth. Average turnover times with respect to sinking were 4.7&#65533;10.9 d, equating to minimum in situ population growth rates of ~ 0.1&#65533;0.2 d&#8722;1. Using simultaneous measurements of sinking organic carbon, we find that these organisms could only meet their carbon demand if their carbon : volume ratio were ~ 1 &#956;g C mm&#8722;3. This value is substantially lower than previously used in global estimates of rhizarian biomass, but is reasonable for organisms that use large siliceous tests to inflate their cross&#8208;sectional area without a concomitant increase in biomass. We found that Aulosphaeridae alone can intercept > 20% of sinking particles produced in the euphotic zone before these particles reach a depth of 300&#8201;m. Our results suggest that the local (and likely global) carbon biomass of Aulosphaeridae, and probably the large Rhizaria overall, needs to be revised downwards, but that these organisms nevertheless play a major role in carbon flux attenuation in the twilight zone.

Abstract: The Costa Rica Dome (CRD) is an open-ocean upwelling ecosystem, with high biomasses of picophytoplankton (especially Synechococcus), mesozooplankton, and higher trophic levels. To elucidate the food web pathways supporting the trophic structure and carbon export in this unique ecosystem, we used Markov Chain Monte Carlo techniques to assimilate data from four independent realizations of &#948;15N and planktonic rate measurements from the CRD into steady state, multicompartment ecosystem box models (linear inverse models). Model results present well-constrained snapshots of ecosystem nitrogen and stable isotope fluxes. New production is supported by upwelled nitrate, not nitrogen fixation. Protistivory (rather than herbivory) was the most important feeding mode for mesozooplankton, which rely heavily on microzooplankton prey. Mesozooplankton play a central role in vertical nitrogen export, primarily through active transport of nitrogen consumed in the surface layer and excreted at depth, which comprised an average 36-46% of total export. Detritus or aggregate feeding is also an important mode of resource acquisition by mesozooplankton and regeneration of nutrients within the euphotic zone. As a consequence, the ratio of passively sinking particle export to phytoplankton production is very low in the CRD. Comparisons to similar models constrained with data from the nearby equatorial Pacific demonstrate that the dominant role of vertical migrators to the biological pump is a unique feature of the CRD. However, both regions show efficient nitrogen transfer from mesozooplankton to higher trophic levels (as expected for regions with large fish, cetacean, and seabird populations) despite the dominance of protists as major grazers of phytoplankton.